Lusus
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Lusus naturae, also known as custodians (also referred to as a custodian), are the Vast Error equivalents to a Guardian. They look after the young trolls.
Cooperi.
The owl lusus is like a owl-creature that Sovara has a custodian. She doesn't live inside of her hive due to its size.
Skulltitan lusus
Dismas' lusus.
Oppossum lusus
Arcjec's lusus.
Chimera lusus
Presumably alive since Zekura was, as they were in the shadows while The Executive was walking through his hometown.
Star Guardian
Albion's lusus.
Lefty
Serpaz's lusus.
Canine lusus
Laivan's lusus.
Ouworm lusus
Occeus' lusus.
Minkmom
Taz's lusus.
Neomacrodon Lusus
Murrit's lusus.
Anglerfish Lusus
Calder's lusus.